The Ecology and Economics of Elm Replacement in Harvard Yard
by Peter Del Tredici, Arnold Arboretum of Harvard University
Return to homepage
Return to Arbor Day
Return to Tree Scavenger Hunt
For more information on elms

This article was from: http://www.arboretum.harvard.edu/LC/Harvyard/Elm_yard.htm

Ulmus americana - 100ft. x 22ft.
Conway, New Hampshire
Photographed in 1930
Before one can answer the question of whether or not to replant the American elm in historic landscapes, one should first ask how this tree came to be so widely planted throughout eastern North America in the first place? The answer can be found in the horticultural literature of the 1800s, the most significant of which is a beautiful book by Lorin Dame and Henry Brooks, published in 1890, Typical Elms and Other Trees
of Massachusetts (see excerpt in Arnoldia 42(2): 49-59, 1982). Because this book was written long before Dutch elm disease had spread to North America, and because it is profusely illustrated, it can be considered a portrait of the species at the pinnacle of its landscape influence. The book also contains important information on the growth rate of the species in landscape situations that can never be recreated. Other important writers from the 1800’s include F. A. Michaux (1819), A. J. Downing (1841), D. J. Browne (1846), F. J. Scott (1870), G. B. Emerson (1875), and C. S. Sargent (1890). In reading over this literature, it is clear that the American elm was widely planted for a number of reasons, only one of which was its great size and beauty. Other, more pragmatic reasons, and can be summarized as follows:

Trunk of the Wethersfield Elm, largest
Elm in the United States in 1924

Dutch Elm Disease
The Dutch elm disease fungus, along with its dispersal agent, the European elm-bark beetle,
arrived in North America in 1930, hidden under the bark of a shipment of European elm burl-logs that had been imported into the United States for the manufacture of veneer (Roueché, 1961). The disease killed American elms so quickly and spread so rapidly that people were at first afraid that the American elm was headed for extinction. Fortunately, this dire prediction has not materialized. The fact is that the species still thrives and reproduces as a wild tree in wet woods and along stream banks throughout eastern North America (Burns and Honkala, 1990). As a landscape plant, however, the American elm is close to extinction. The grand old specimens that once graced virtually every town common in New England, four to five feet in diameter, have been replaced by trees that are seldom more than two feet across.
Since the 1960s, there has been considerable publicity surrounding efforts to "bring back" the American elm. The long-term approach has involved either the selection of DED-resistant American elm cultivars or the hybridization of DED-resistant elm species with one another. In contrast, the short-term approach focuses on the preservation of existing specimens of the American elm and includes a program of spraying for the beetle, injection of infected trees with fungicides, and the vigilant removal of diseased limbs. While such extraordinary treatments have succeeded in saving selected individuals for up to twenty years, the inevitable infirmities of old age are now catching up with older trees. Such a preservation strategy is, at best, a temporary solution.

Case Study: The Elms in Harvard Yard
The American elm has been the main stay of the Harvard Yard landscape for well over a hundred years. While there have been many threats to the integrity of this planting during this time, none of them have been as serious as the introduction of Dutch elm disease. In 1979, when Harvard University began to implement an integrated elm protection program, there were 285 elms on campus. Most of them were American elms, but there were also a

An old American Elm in
Harvard Yard
number of English and European elms of uncertain identity mixed in. Most of the trees had been planted during the early 1900s, and were about seventy or eighty years old when the protection program began. By 1994, after fifteen years of treatment, there were only 165 elms left. This amounts to a 42% mortality. The average cost of the total protection program over this time period has been approximately $25,000 per year, broken down as follows:

Over the 15 year period the total amount spent on the elms was approximately $375,000 or $100 per tree per year. Despite this treatment, there was a 42% mortality at the end of the 15 year period. Extending this figure out, the cost of elm maintenance over a twenty year period would be roughly $500,000 with a mortality approaching 50%. For comparative purposes, the annual cost for maintaining non-elm trees in the Yard is approximately $20 per tree per year.
Replacement Costs
In 1994 an elm replacement program was begun (Shaw, 1994; Van Valkenburgh and Del Tredici, 1994). The cost of replanting two hundred new trees in the Yard, most of which were four to eight inches in caliper and ten to twenty feet tall, was $470,000. This works out to an average of $2,350 per tree planted and watered, including a one year maintenance contract and guarantee.

A mixed planting of old and and new trees in
Harvard Yard
Essentially the numbers indicate that the cost of planting two hundred new trees was roughly equal to cost of maintaining two hundred and eighty-five elms for twenty years, after which time only half of them will still be alive and the other half will be in a state of decline. To put it another way, twenty years of maintenance of one large elm, with only a 50% chance of survival, equals the cost of planting one new four to eight inch caliper tree.
There is no absolute answer to the question of how much one should invest in an elm protection program. All that can be said with certainty is that any elm protection program should always be undertaken in conjunction with a tree planting program of species other than elms. Planting new trees is an investment in the future that softens the blow when a big elm inevitably dies.
Achieving Diversity
Unfortunately very few, if any, trees have the same combination of graceful form and great size that the American elm does. The honey locust, Gleditsia triacanthos, comes about as close as any tree, but is much slower growing. Zelkova serrata has roughly the same shape, but is much smaller. As Koller and Weaver (1982) point out, there is no exact replacement for the American elm. The key to successful substitution is to look for species that have the same landscape impact or stature as the American elm, not necessarily the same structure.
By working within a given genus or family, one can achieve diversity and uniformity simultaneously. In the Tercentenary Theater part of the Yard, for example, we chose to group various legumes together which all share the characteristic arching trunk and

A classic Harvard combination:
Bricks and Elms
flat-topped crown, but are clearly different in other aspects of their habit. These include: Gleditsia, Sophora, Gymnocladus, and Cladrastis. Similarly, one can also group different oaks together to achieve a measure of uniformity amidst diversity. In the oldest section of the Yard, we used a variety of Quercus species, including rubra, palustris, phellos, coccinea, alba, bicolor, and acutissima.
As regards the genus Ulmus itself, it must be remembered that not all elms are equal. Just because a tree is an elm does not mean it can be used as a replacement for Ulmus americana. The Siberian elm (Ulmus pumila), for example, is highly resistant to DED but is very messy and quite graceless in form. The lacebark elm (Ulmus parviflora) is a handsome tree, but much smaller in stature than its American cousin. Some of the more recent hybrid elms may eventually turn out to be excellent replacements, but they have not yet been thoroughly tested (Dirr, 1990; Santamour and Bentz, 1995; Townsend et al., 1995).
It must also be remembered that there are several other diseases besides DED that can kill American elms (Sinclair et al., 1987). In particular, phloem necrosis and elm yellows can be lethal to many of the elms that have been selected for their resistance to DED (Santamour and Bentz, 1995). It should also be remembered that the elm leaf beetle, along with a host of other insects, had been decimating elm plantings long before DED arrived on the scene. If the American elm is to make a return to the modern American landscape-be it either as a hybrid or as a disease resistant selection-then it should be on an equal footing with other trees, never as an all encompassing landscape theme.
One can summarize the advantages to increasing landscape diversity can be summarized as follows:

Conclusions
The desire to bring back the American elm is basically a romantic (i. e. emotional) ideal. Granting, for the moment, the validity of the desire to bring the tree back, one still

Ulmus americana - Chelmsford Center, Massachusetts
Photographed in 1925
needs to ask, "Bring it back at what density?" Clearly the high density of American elms in many cities and towns that existed during the first third of the twentieth century (Dawson and Khawaja, 1985) is not a situation that ought to be recreated. Indeed, it was the high density of American elms that allowed the elm bark beetle population to build up rapidly. leading to the epidemic spread of DED. One sees more elms surviving these days

Ulmus americana - 90ft. x 10ft.
Mansfield, Massachusetts
Photographed in 1925
than in the past not because trees are more resistant to Dutch Elm Disease than they used to be, but because the reduced elm population has resulted in lower elm bark beetle populations. This, in turn, allows more elms to escape detection by their predators.
If "restoring" a given historic landscape means replanting the American elm-or any of its resistant selections or hybrids-to the density it occupied historically, then that is a mistake. In the popular literature on elms, the unspoken assumption always seems to be that if one could only solve the disease resistance problem, then one could recreate the grand, elm-lined streets of the past. This idea is biologically unsound. Because of the dynamic nature of the interaction between host and predator, however, disease resistance must always be considered a relative phenomenon-always in flux-rather than a fixed genetic trait. Historical accuracy not withstanding, it is not in anyone’s interest to bring the American elm back at its former position of landscape preeminence.

References
Browne, D. J. 1846. Trees of America. Harper & Bros., New York.
Burns, R. M. and B. H. Honkala, eds. 1990. Ulmus americana, in Silvics of North America, vol. 2,
hardwoods. U. S. D. A. Forest Service Agriculture Handbook 654.
Dame, L. L. and H. Brooks. 1890. Typical Elms and Other Trees of Massachusetts. Little Brown and
Co., Boston.
Dawson, J. O. and M. A. Khawaja. 1985. Change in street-tree composition of two Urbana, Illinois
neighborhoods after fifty years: 1932-1982. Journal of Arboriculture 11(11): 344-348.
Dirr, M. 1990. Manual of Woody Landscape Plants, fourth ed. Stipes Publ. Co., Champaign, Illinois.
Downing, A. J. 1875. A Treatise on the Theory and Practice of Landscape Gardening, ninth ed.
Orange Judd, Co., New York.
Emerson, G. B. 1875. A Report on the Trees and Shrubs Growing Naturally in the Forests of
Massachusetts, 2nd ed. Little Brown and Co., Boston
Koller, G. L. and R. E. Weaver, Jr. 1982. Replacing the American elm: twelve stately trees. Arnoldia
42(2): 88-100.
Michaux, F. A. 1819. Ulmus americana, in The North American Sylva, vol. 3, translated by A. L.
Hillhouse. Paris and Philadelphia.
Roueché, B. 1961. Profiles: A great green cloud. The New Yorker July 15, 1961: 35-53.
Santamour, F. S., Jr. and S. E. Bentz. 1995. Updated checklist of elm (Ulmus) cultivars for use in North
America. J. Arboriculture 21(3): 122-131.
Sargent, C. S. 1890. Ulmus americana, in Silva of North America, vol. 7. Houghton and Mifflin,
Boston.
Scott. F. J. 1870. The Art of Beautifying Suburban Home Grounds of Small Extent. D. Appleton,
New York.
Shaw, J. 1994. Every tree doomed. Harvard Magazine 96(4): 46-53.
Sinclair, W. A., H. H. Lyon, and W. T. Johnson. 1987. Diseases of Trees and Shrubs. Cornell U. Press,
Ithaca, New York.
Townsend, A. M. , S. E. Bentz, and G. R. Johnson. 1995. Variation in response of selected American elm
clones to Ophiostoma ulmi. Journal of Environmental Horticulture 13(3): 126-128.
Van Valkenburgh, M. and P. Del Tredici. 1994. Restoring the Harvard Yard Landscape. Arnoldia 54 (1):
2-11.

Return to homepage
Return to Arbor Day
Return to Tree Scavenger Hunt
For more information on elms